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Sibon canopy ARTEAGA & BATISTA, 2023

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common Names 
SynonymSibon canopy ARTEAGA & BATISTA 2023: 16 
DistributionPanama

Type locality: Cerro Gaital, La Pintada, Coclé province, Panama (8.70874, -80.42411; 543 m a.s.l.).  
Reproduction 
TypesHolotype: MHCH 3110 (Figs 5b, 7), adult female collected by Abel Batista on 8 August 2016 at Cerro Gaital, La Pintada, Coclé province, Panama (8.70874, -80.42411; 543 m a.s.l.).
Paratypes: MHCH 1067, SMF 88713–14, juveniles collected by Johannes Köhler, Abel Batista, and Marcos Ponce on 17 January 2007 at Casa de Ancón, Sendero el Pianista, Bocas del Toro province, Panama (8.87142, -82.41594; 1005 m a.s.l.). MHCH 220, juvenile female collected by Abel Batista and Marcos Ponce on March 2002 at Camino al Río Culebra, Bocas del Toro province, Panama (8.90772, -82.39115; 698 m a.s.l.). SMF 85077, adult female collected by Gunther Köhler, Abel Batista, Marcos Ponce, and Javier Sunyer on 17 January 2006 at Reserva Forestal La Fortuna, Comarca Ngäbe-Buglé, Panama (8.77763, -82.20916; 1030 m a.s.l.). SMF 89596, adult female collected by Leonhard Stadler and Nadim Hamad on 5 August 2008 at Cerro Mariposa, Veraguas province, Panama (8.52488, -81.13275; 679 m a.s.l.). SMF 90023, adult female collected by Arcadio Carrizo on 27 June 2008 at Cerro Negro, Veraguas province, Panama (8.56901, -81.09894; 680 m a.s.l.). SMF 91578, adult female collected by Sebastian Lotzkat and Andreas Hertz on 17 July 2010 at Río Changena, Bocas del Toro province, Panama (8.97851, -82.69005; 1641 m a.s.l.). SMF 86411, juvenile collected by Abel Batista and Marcos Ponce on 10 February 2006 at Sendero El Pianista, Bocas del Toro province, Panama (8.87141, -82.41594; 1005 m a.s.l.). SMF 90208, juvenile collected by Joe-Felix Bienentreu and Frank Hauenschild on 25 October 2009 at Cerro Guayabo, Chiriquí province, Panama (8.75531, -82.25431; 1247 m a.s.l.). MHCH 2363–64, males collected by Sebastian Lotzkat and Andreas Hertz between 29 October 2009 and 11 June 2010 at Cabeceras del Río Chiriquí Mali, Comarca Ngäbe-Buglé, Panama (8.78906, -82.21547; 1080 m a.s.l.). MHCH 2365, juvenile male collected by Sebastian Lotzkat and Andreas Hertz on 7 August
2010 at Cerro Mariposa, Veraguas province, Panama (8.50815, -81.12104; 899 m a.s.l.). SMF 85078, adult male collected by Gunther Köhler, Abel Batista, Marcos Ponce, and Javier Sunyer on 19 January 2006 at Reserva Forestal La Fortuna, Comarca Ngäbe-Buglé, Panama (8.77763, -82.20916; 1030 m a.s.l.). SMF 88715, adult male collected by Sebastian Lotzkat and Andreas Hertz on 14 May 2008 at Trail to Rio Hornito, Chiriquí province, Panama (8.67385, -82.21845; 1320 m a.s.l.). SMF 89597, adult male collected by Leonhard Stadler and Nadim Hamad on 6 August 2008 at Cerro Mariposa, Veraguas province, Panama (8.51463, -81.11927; 1003 m a.s.l.). SMF 89786, adult male collected by Sebastian Lotzkat and Andreas Hertz on 1 April 2009 at Cerro Negro, Veraguas province, Panama (8.56901, -81.09894; 900 m a.s.l.). SMF 90024, adult male collected by Arcadio Carrizo on 29 July 2008 at Cerro Negro, Veraguas province, Panama (8.57697, -81.09705; 1085 m a.s.l.). SMF 90207, adult male collected by Sebastian Lotzkat and Andreas Hertz on 29 October 2009 at Cabeceras del Río Chiriquí Mali, Comarca Ngäbe-Buglé, Panama (8.78906, -82.21547; 1054 m a.s.l.). SMF 91579, adult male collected by Sebastian Lotzkat and Andreas Hertz on 11 June 2010 at Cabeceras del Río Chiriquí Mali, Comarca Ngäbe-Buglé, Panama (8.78906, -82.21547; 1054 m a.s.l.). SMF 91580, adult male collected by Sebastian Lotzkat and Andreas Hertz on 24 June 2010 at Bosque Guayabito, Comarca Ngäbe-Buglé, Panama (8.54939, -81.48467; 1510 m a.s.l.). 
DiagnosisDiagnosis: Sibon canopy sp. nov. is placed in the genus Sibon based on phylogenetic evidence and on having the penultimate supralabial conspicuously higher than all other supralabials. The species is diagnosed based on the following combination of characters: (1) 15/15/15 smooth dorsals with enlarged vertebral row (1.4× as wide as adjacent rows); (2) loreal and prefrontal in contact with orbit; (3) 7–8 supralabials with, usually, 4th, 5th, and occasionally 6th contacting orbit; (4) usually 7–8 infralabials with 2nd–6th in contact with chinshields, first pair of infralabials not in contact behind symphysial due to presence of a postmental; (5) 180–189 ventrals in males, 170–185 in females; (6) 113–130 divided subcaudals in males, 107–124 in females; (7) dorsal background color olive with maroon bands (1–2 dorsal scales long mid-dorsally and 3–5 dorsal scales long on the lower flanks) and a reddish tint along the vertebral line, ventral surfaces white with encroachment from the dorsal maroon blotches, dorsal aspect of head composed of broad irregular maroon to blackish blotches interspersed with olive to red blotches, throat white with brownish blotches, iris dark reddish brown; (8) 336–427 mm SVL in males, 318–357 mm in females; (9) 160–221 mm TL in males, 157–185 mm in females (Arteaga & Batista 2023). 
CommentDiet: Ray et al. (2012) found oligochaete and mollusk remains in fecal samples of 37 individuals of S. canopy from El Copé and Altos del María, Panama. They also observed an individual feeding on a snail at El Copé. 
EtymologyThe specific epithet canopy is used as a noun in apposition and honors the Canopy Family system of reserves, particularly its Canopy Lodge in Valle de Antón, Coclé province, Panama, where the new species occurs. 
References
  • Arteaga, A., & Batista, A. 2023.  A consolidated phylogeny of snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Colombia, Ecuador, and Panama. ZooKeys, 1143: 1-49 - get paper here
 
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