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Ptyas doriae (BOULENGER, 1888)

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Doria's Green Snake
E: Hampton's Green Snake [hamptoni]
Chinese: 纯绿翠青蛇 
SynonymCyclophiops doriae BOULENGER 1888: 599
Ablabes doriae — BOULENGER 1890
Ablabes hamptoni BOULENGER 1900
Liopeltis hamptoni — WALL 1924: 865
Liopeltis doriae — WALL 1924
Eurypholis doriae — POPE 1935: 281
Opheodrys hamptoni — SMITH 1943: 180
Entechinus doriae — CUNDALL 1981
Liopeltis doriae — WELCH 1988
Entechinus doriae — WELCH 1988
Entechinus doriae — MCDOWELL & JENNER 1988
Entechinus hamptoni — MCDOWELL & JENNER 1988
Cyclophiops hamptoni — TORIBA 1989
Cyclophiops doriae — ZHAO & ADLER 1993
Ophiodrys doriae — DAS 1996: 58
Opheodrys doriae — SHARMA 2004
Cyclophiops doriae — ZHAO 2006: 189
Cyclophiops doriae — WALLACH et al. 2014: 202
Cyclophiops hamptoni — WALLACH et al. 2014: 202
Ptyas hamptoni — FIGUEROA et al. 2016
Ptyas doriae — FIGUEROA et al. 2016
Cyclophiops doriae — WANG et al. 2022 
DistributionChina (Yunnan), Myanmar (= Burma) (Kachin Hills), India (Assam)

Type locality: “Monti ad Est di Bhamò: Kakhien Hills (Birmania)” [= mountains to the east of Bhamò (24°15’N, 97°14’E), ext. SE Kachin Division, Myanmar]”

hamptoni: Myanmar (= Burma); Type locality: Mogok, Burma  
Reproductionoviparous 
TypesLectotype: MSNG 30384a, adult male; collected by L. Fea in 1886. Lectotype designated by Capocaccia (1961). Paralectotype: (n=1) MSNG 30384B; same information as the holotype.
Holotype: BMNH 1900.9.20.15; BMNH 1946. 1.5.32 fide Meetei et al. 2018 [hamptoni] 
Diagnosis 
CommentType species: Cyclophiops doriae is the type species of the genus Cyclophiops Boulenger 1888.

Synonymy: partly after SMITH 1943; Figueroa et al. 2016: 21 and Li et al. 2020 synonymized Cyclophiops with Ptyas. Meetei et al. 2018 synonymized hamptoni with doriae.

Abundance: both P. doriae and P. hamptoni are very rare; P. hamptoni is known only from the type, which is a female. There seem to be no photos of live P. doriae. It is still possible they are different species.

Habitat: partly arboreal (Harrington et al. 2018). 
EtymologyNamed after Marchese Giacomo Doria (1840-1913), Italian zoologist who collected in Persia (Iran) with de Filippi (1862-1863) and in Borneo with Beccari (1865-1866). He founded the Museo Civico di Storia Naturale, Turin (1867-1913), and was its first Director. 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Boulenger, G.A. [spelled as Boulanger] 1900. Description of a new Snake of the genus Ablabes from Burma. J. Bombay Nat. Hist. Soc. 13: 553 - get paper here
  • Boulenger, GEORGE A. 1888. An account of the Reptilia obtained in Burma, north of Tenasserim, by M. L. Fea, of the Genova Civic Museum. Ann. Mus. Civ. Stor. Nat. Genova, ser. 2, 6: 593-604 - get paper here
  • Boulenger, George A. 1890. The Fauna of British India, Including Ceylon and Burma. Reptilia and Batrachia. Taylor & Francis, London, xviii, 541 pp. - get paper here
  • Boulenger, George A. 1894. Catalogue of the snakes in the British Museum (Natural History). Volume II., Containing the Conclusion of the Colubridæ Aglyphæ. British Mus. (Nat. Hist.), London, xi, 382 pp. - get paper here
  • Boulenger,G.A. 1900. Description of a new snake of the genus Ablabes from Burma. Ann. Mag. Nat. Hist. (7) 6: 409 - get paper here
  • Capocaccia, L. 1961. Catalogo dei tipi di Rettili del Museo Civico di Storia Naturale di Genova [MSNG]. Ann. Mus. Civ. Stor. Nat. Giacomo Doria 72: 86-111 - get paper here
  • Dowling, H.G., & Jenner, J.V. 1988. Snakes of Burma: checklist of reported species and bibliography. Smithsonian Herp. Inf. Serv. (76): 19 pp. - get paper here
  • Figueroa A, McKelvy AD, Grismer LL, Bell CD, Lailvaux SP 2016. A Species-Level Phylogeny of Extant Snakes with Description of a New Colubrid Subfamily and Genus. PLoS One 11 (9): e0161070.doi:10.1371/journal.pone.0161070 - get paper here
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Li, J-N., Liang, D., Wang, Y-Y., Guo, P., Huang, S., Zhang, P. 2020. A Large-scale Systematic Framework of Chinese Snakes Based on a Unified Multilocus Marker System. Molecular Phylogenetics and Evolution - get paper here
  • Sclater,W.L. 1891. Notes on a collection of snakes in the Indian Museum, with descriptions of several new species. J. Asiat. Soc. Bengal LX: 230-250 - get paper here
  • Sharma, R. C. 2004. Handbook Indian Snakes. AKHIL BOOKS, New Delhi, 292 pp.
  • Smith, M.A. 1943. The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. 3 (Serpentes). Taylor and Francis, London. 583 pp.
  • Toriba, M. 1989. A book review: Snake of the Orient, a Checklist, by Kenneth R. G. Welch, 1988, Krieger, Melbourune, Florida, vii+183pp. (in Japanese). Japanese Journal of Herpetology 13 (1): 21-25.
  • Wall,F. 1924. A Hand-list of the Snakes of the Indian Empire. Part 3. J. Bombay nat. Hist. Soc. 29: 864-878 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Wang K, Lyu ZT, Wang J, Qi S, Che J 2022. Updated Checklist and Zoogeographic Division of the Reptilian Fauna of Yunnan Province, China. Biodiversity Science 30 (4): 21326, 1–31 - get paper here
  • Wang, Kai; Jinlong Ren, Hongman Chen, Zhitong Lyu, Xianguang Guo Ke Jiang, Jinmin Chen, Jiatang Li, Peng Guo, Yingyong Wang, Jing Che 2020. The updated checklists of amphibians and reptiles of China. Biodiversity Science 28 (2): 189-218 - get paper here
  • Zhao, E.M. 2006. The snakes of China [in Chinese]. Hefei, China, Anhui Sience & Technology Publ. House, Vol. I, 372 pp., Vol. II (color plates), 280 pp.
 
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