Philodryas arnaldoi (AMARAL, 1932)

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Higher Taxa	Colubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies
Common Names	E: Arnaldo's Green Racer Portuguese: Papa-Pinto, Papa-Rato, Parelheira-Clara, Parelheira-do-Mato
Synonym	Chlorosoma arnaldoi AMARAL 1932: 100 Philodryas arnaldoi — GRAZZIOTIN et al. 2012 Philodryas arnaldoi — WALLACH et al. 2014: 549 Philodryas arnaldoi — NOGUEIRA et al. 2019 Philodryas arnaldoi — ARREDONDO et al. 2020 Pseudablabes arnaldoi — MELO-SAMPAIO et al. 2020
Distribution	Brazil (Sao Paulo, Parana, Rio Grande do Sul, Santa Catarina) Type locality: S. Bento, Santa Catarina, Brazil.
Reproduction	oviparous
Types	Holotype: IBSP (= IB) 7807, female
Diagnosis
Comment	Habitat: fully arboreal (Harrington et al. 2018). Phylogenetics: MELO-SAMPAIO et al. 2020 tentatively included arnaldoi within Pseudablabes (based on morphology) although they did not have any molecular data. Note that Arredondo et al. 2020 could not replicate the monophyly of Pseudablabes as defined by Melo-Sampaio et al., hence we reverted them to Philodryas.
Etymology	Named after Arnaldo França, Amaral's technical assistant.
References	Amaral,A. do 1932. Contribuicao ao conhecimento dos ofidios do Brasil VI. Uma nova especie de Colubrideo opisthoglypho, do genero Chlorosoma Wagler, 1830. Mem. Inst. Butantan 7: 99-101 + 1 plate - get paper here Arredondo, J. C., Grazziotin, F. G., Manfrini, G. J. S., Rodrigues, M. T. U., Bonatto, S. L., & Zaher, H. E. D. 2020. Molecular phylogeny of the tribe Philodryadini Cope, 1886 (Dipsadidae: Xenodontinae): Rediscovering the diversity of the South American Racers. Papéis Avulsos de Zoologia, 60, e20206053-e20206053 - get paper here Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here Bressan, Paulo Magalhães; Maria Cecília Martins Kierulﬀ, Angélica Midori Sugieda (eds.) 2009. FAUNA AMEAÇADA DE EXTINÇÃO NO ESTADO DE SÃO PAULO: VERTEBRADOS. São Paulo: Fundação Parque Zoológico de São Paulo: Secretaria do Meio Ambiente, 2009. - get paper here Gonzalez R. C. et al. 2020. Lista dos Nomes Populares dos Répteis no Brasil – Primeira Versão. Herpetologia Brasileira 9 (2): 121 – 214 - get paper here Grazziotin, Felipe G.; Hussam Zaher, Robert W. Murphy, Gustavo Scrocchi, Marco A. Benavides, Ya-Ping Zhang and Sandro L. Bonatto 2012. Molecular phylogeny of the New World Dipsadidae (Serpentes: Colubroidea): a reappraisal. Cladistics DOI: 10.1111/j.1096-0031.2012.00393.x - get paper here Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here Kunz, Tobias Saraiva;<br />Ivo Rohling Ghizoni-Jr.,<br />Walter Luis Alves dos Santos & Paulo Afonso Hartmann 2007. Nota sobre a coleção herpetológica da Universidade Federal de Santa Catarina (UFSC). Biotemas, 20 (3): 127-132 - get paper here Lema, Thales de 1962. Ocorréncia de Philodryas arnaldoi (Amaral, 1932) no estado do Rio Grande do Sul, Brasil (Serpentes, Colubridae). Iheringia, Série Zoologia 22: 1-4 + 2 plates MARQUES, O. A. V. et al. 2009. Repteis. In: Bressan et al., FAUNA AMEAÇADA DE EXTINÇÃO NO ESTADO DE SÃO PAULO: VERTEBRADOS. São Paulo: Fundação Parque Zoológico de São Paulo: Secretaria do Meio Ambiente, pp. 285-327 - get paper here Melo-Sampaio, P. R., Passos, P., Martins, A. R., Moura-leite, J. C., Morato, S. A., Venegas, P. J., Chávez G., Venâncio N.M. & De Souza, M. B. 2020. A phantom on the trees: integrative taxonomy supports a reappraisal of rear-fanged snakes classification (Dipsadidae: Philodryadini). Zoologischer Anzeiger 290: 19-39 [dated 2021 in print version, but 2020 published online] - get paper here Morato, Sérgio Augusto Abrahão; Renato Silveira Bérnils, Julio Cesar de Moura-Leite 2017. RÉPTEIS DE CURITIBA: COLETÂNEA DE REGISTROS Hori Consultoria, Curitiba, Paraná, Brasil, 99 pp. Nogueira, Cristiano C.; Antonio J.S. Argôlo, Vanesa Arzamendia, Josué A. Azevedo, Fausto E. Barbo, Renato S. Bérnils, Bruna E. Bolochio, Marcio Borges-Martins, Marcela Brasil-Godinho, Henrique Braz, Marcus A. Buononato, Diego F. Cisneros-Heredia, 2019. Atlas of Brazilian snakes: verified point-locality maps to mitigate the Wallacean shortfall in a megadiverse snake fauna. South American J. Herp. 14 (Special Issue 1):1-274 - get paper here Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp. Zaher, Hussam; Fausto Erritto BarboI; Paola Sanchez Martínez; Cristiano Nogueira; Miguel Trefaut Rodrigues; Ricardo Jannini Sawaya 2011. Répteis do Estado de São Paulo: conhecimento atual e perspectivas. Biota Neotropica, 11 (1): 1–15. - get paper here
External links	IUCN Red List http://www.pucrs.br/~museu/http/pro-mata/chavserp.htm http://www.lacerta.de/AS/Taxon.php?Genus=32&Species=114 Google images

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