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Lycodon pictus JANSSEN, PHAM, NGO, LE, NGUYEN & ZIEGLER, 2019

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common Names 
SynonymLycodon pictus JANSSEN, PHAM, NGO, LE, NGUYEN & ZIEGLER 2019 
DistributionN Vietnam (Cao Bang Province), China (Guangxi Zhuang)

Type locality: Trung Khanh District, Cao Bang Province, (altitude 701 m a.s.l.)  
Reproduction 
TypesHolotype. IEBR 4166 (field number CB 2012.97), adult male, collected on 18 April 2012 by TQN et al. Paratypes. ZFMK 93747, juvenile, collected on 15 October 2011 by TQN et al. (altitude 588 m a.s.l.), Ha Lang District, Cao Bang Province; ZFMK 93746, adult fe- male, collected on 10 April 2012 by TQN et al., Ha Lang District, Cao Bang Province. 
DiagnosisDiagnosis: Lycodon pictus sp. nov. can be differentiated from its congeners by the following morphological characters: dorsal scales in 17–17–15 rows, all smooth; su- pralabials usually eight (rarely nine); infralabials ten; one elongated loreal on each side, in contact with the eye; precloacal plate single; ventral scales 212–218 (plus one or two preventral scales); subcaudals 90 or 91; a total length of 597+ mm in males and 543 mm in females; tail / total length ratio 0.211–0.215; maxillary teeth 13 or 14; dorsal surface of body with 28 or 29 light body bands; dorsal surface of tail with 13 cream bands forming a distinct blotch in the vertebral region; ventral surface of body and tail mostly cream with the dark body bands in part extending towards the venter, some- times forming complete dark bands around the body. 
CommentHabitat: karst environment. Specimens were found at night between 19:00 and 23:00, on forest paths or on the ground near cave entrances. The surrounding habitat was secondary karst forest, consisting of medium and small hardwood trees mixed with shrubs and vines. Air temperature was 23.4–29.6°C and humidity was 66–79%.

Sympatry: Acanthosaura lepidogaster, Gekko adleri, Goniurosaurus luii, Lycodon futsingensis, Protobothrops trungkhanhensis. 
EtymologyThe name of the species pictus means painted or decorated in Latin and refers to its unique dorsal colour pattern. 
References
  • Janssen HY, Pham CT, Ngo HT, Le MD, Nguyen TQ, Ziegler T 2019. A new species of Lycodon Boie, 1826 (Serpentes, Colubridae) from northern Vietnam. ZooKeys 875: 1-29 - get paper here
  • Janssen, H. Y., Jin-Long, R., Jia-Tang, L., Zeng, W., Nguyen, T. T., Nguyen, T. Q., ... & Ziegler, T. 2020. Range extension and extended diagnosis of Lycodon pictus: First country record from China. Revue suisse de Zoologie, 127(2), 413-422 - get paper here
  • Kwet, A. 2020. Liste der im Jahr 2019 neubeschriebenen Reptilien. Elaphe 2020 (3): 44-67
 
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