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Lycodon chapaensis (ANGEL & BOURRET, 1933)

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Namdong Wolf Snake
G: Namdong Wolfszahnnatter
Vietnamese: Rắn khuyết nam động 
SynonymDinodon septentrionale chapaense ANGEL & BOURRET 1933: 131
Dinodon septentrionale chapaense — BOURRET 1935: 6
Dinodon septentrionale var. chapaense — DEUVE 1961: 16
Dinodon septentrionale — ZHAO & YANG 1997
Dinodon septentrionale — ZHAO et al. 1998
Dinodon septentrionale — HE & ZHOU 2002
Dinodon septentrionale — ZHAO, 2006
Dinodon septentrionale — YANG & RAO 2008
Lycodon septentrionalis — SILER et al. 2013
Lycodon septentrionalis — GUO et al. 2013
Lycodon septentrionalis — CAI et al. 2015 (in part)
Lycodon septentrionalis — JIANG et al. 2016
Lycodon cf. septentrionalis — YANG et al. 2019
Lycodon namdongensis LUU, ZIEGLER, HA, LE & HOANG 2019
Lycodon septentrionalis — WANG et al., 2020
Lycodon chapaensis — WANG et al. 2020 
DistributionVietnam (Thanh Hoa), China (Yunnan)

Type locality: 20 km SW of Lao-Kay (=Lao Cai), Tonkin, Vietnam

namdongensis: Type locality: karst forest, Nam Dong Nature Reserve, Quan Son District, Thanh Hoa Province, Vietnam (20°18.298’N; 104°54.776’E, at an elevation of 616 m a.s.l.)  
Reproduction 
TypesHolotype: MNHN-RA-1933.0011, adult female. Collected by Bourret R. on 01 July 1931.
Holotype: VNUF R.2017.23 (field number: TT 17.23), adult male, collected on 13 June 2017 by Vinh Quang Luu, Nghia Van Ha, Oanh Van Lo, and Ngoan Van Ha [namdongensis] 
DiagnosisDiagnosis: Lycodon namdongensis sp. nov is characterized by the following morphological characters: dorsal scales in 17−17−15 rows, all smooth; supralabials 8; infralabials 10; one large loreal on each side, separated from the eye; cloacal plate single; ventral scales 218 (plus two preventral scales); subcaudals 85; a total length of 723 mm in the single male; tail / total length ratio 0.205; dorsal head pattern whitish grey; dorsal surface of body with 23 narrowed greyish cream bands; dorsal surface of tail with 14 cream bands; ventral surface of body mostly cream with a few dark spots posteriorly; ventral surface of tail dark grey. 
CommentHabitat: ground near shrubs in secondary karst forest.

NCBI taxonID: 2546291 [namdongensis] 
EtymologyThe species name is derived from the type locality of this species (Chapa, = Sa Pa, in Lao Cai Province of Vietnam). 
References
  • Angel, F. & BOURRET,R. 1933. Sur une pe tite collection de serpents du Tonkin. Descriptions d'espèce nouvelles. Bull. Soc. Zool. Fr. 58: 129-140
  • Bourret,R. 1935. Notes herpétologiques sur l'Indochine Française X. Les serpents de la station d’altitude du Tam-dao. Bull. Gen. Instr. Pub. Hanoi 8 (avril): 259-271 (1-13)
  • Janssen HY, Pham CT, Ngo HT, Le MD, Nguyen TQ, Ziegler T 2019. A new species of Lycodon Boie, 1826 (Serpentes, Colubridae) from northern Vietnam. ZooKeys 875: 1-29 - get paper here
  • LUU, VINH QUANG; THOMAS ZIEGLER, NGHIA VAN HA, MINH DUC LE, TUOI THI HOANG 2019. A new species of Lycodon Boie, 1826 (Serpentes: Colubridae) from Thanh Hoa Province, Vietnam. Zootaxa 4586 (2): 261-277 - get paper here
  • Wang K, Lyu ZT, Wang J, Qi S, Che J 2022. Updated Checklist and Zoogeographic Division of the Reptilian Fauna of Yunnan Province, China. Biodiversity Science 30 (4): 21326, 1–31 - get paper here
  • Wang K, Yu ZB, Vogel G, Che J. 2020. Contribution to the taxonomy of the genus Lycodon H. Boie in Fitzinger, 1827 (Reptilia: Squamata: Colubridae) in China, with description of two new species and resurrection and elevation of Dinodon septentrionale chapaense Angel, Bourret, 1933. Zool Res 42 (1): 62-86 [online 2020, in print 2021] - get paper here
 
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