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Hebius maximus (MALNATE, 1962)

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Higher TaxaColubridae (Natricinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Western China Keelback
Chinese: 华西腹链蛇 (Huá Xī Fù Liàn Shé) 
SynonymAmphiesma sauteri maximus MALNATE 1962: 272
Natrix sauteri — POPE 1935: 125 (in part)
Natrix sauteri — WU 1985: 209 (in part)
Amphiesma sauteri — INGER et al. 1990: 20
Amphiesma sauteri — ZHAO & ADLER 1993: 228 (in part)
Amphiesma sauteri — ZHAO et al. 1998: 80 (in part)
Amphiesma sauteri — ZHAO 2003: 140
Amphiesma sauteri — DAVID et al. 2007: 60 (in part)
Amphiesma sauteri — WALLACH et al. 2014: 33 (in part).
Hebius sauteri — GUO et al. 2014: 428 (in part)
Hebius sauteri — DAVID et al. 2021 (in part)
Hebius maximus — LI et al. 2022 
DistributionSW China (Chongqing: Youyang Tujia and Miao Autonomous County, Xiushan Tujia and Miao Autonomous County, Sichuan: Mabian County, Pingshan County, Yingjing County, Hongya County, Dujiangyan City, Mianyang City, Leshan City, Guizhou: Qixingguan District, Bijie City)

Type locality: Hsaioyangchi, Szechwan, China (= Xiaoyangxi, Mabian Yi Autonomous County, Leshan City, Sichuan Province, China)  
Reproductionoviparous 
TypesHolotype: FMNH 18796 (formerly CHNM 18796), a 457 mm adult female specimen (Fig. 7). Paratypes. FMNH 18797, USNM 71570, USNM 84362. 
Diagnosis 
CommentSynonymy: after Li et al. 2022, who suggested to elevate maximus to full species level, first in a preprint, then in a peer-reviewed paper. 
EtymologyHebius maximus comb. nov. is named for its highest count of ventral scales among three previous subspecies of H. sauteri (Malnate 1962). 
References
  • David, P., Bain, R. H., Nguyen Quang Truong, Orlov, N. L., Vogel, G., Vu Ngoc Thanh & T. Ziegler 2007. A new species of the natricine snake genus Amphiesma from the Indochinese Region (Squamata: Colubridae: Natricinae). Zootaxa 1462: 41-60 - get paper here
  • DAVID, PATRICK; GERNOT VOGEL, TRUONG QUANG NGUYEN, NIKOLAI L. ORLOV, OLIVIER S. G. PAUWELS, ALEXANDRE TEYNIÉ, THOMAS ZIEGLER 2021. A revision of the dark-bellied, stream-dwelling snakes of the genus Hebius (Reptilia: Squamata: Natricidae) with the description of a new species from China, Vietnam and Thailand. Zootaxa 4911: 1–61 - get paper here
  • Inger R F; Zhao E; Bradley Shaffer H; Guanfu Wu 1990. Report on a collection of amphibians and reptiles from Sichuan, China. Fieldiana: Zoology (No. 58) : i-iii, 1-24 - get paper here
  • Li M-L, Ren J-L, Huang J-J, Lyu Z-T, Qi S, Jiang K, Wang Y-Y, Li J-T 2022. On the validity of Hebius sauteri maximus (Malnate, 1962) (Squamata, Natricidae), with the redescription of H. maximus comb. nov. and H. sauteri (Boulenger, 1909). Herpetozoa 35: 265-282 - get paper here
  • Li, M.L., Ren, J.L., Huang, J.J., Lyu, Z.T., Qi, S., Jiang, K., Wang, Y.Y. and Li, J.T., 2022. On the validity of Hebius sauteri maximus (Squamata, Natricidae), with redescription of H. maximus comb. nov. and H. sauteri from China. ARPHA Preprints, 3, p.e94937 - get paper here
  • Malnate, E.V. 1962. The relationships of five species of the Asiatic natricine snake genus Amphiesma. Proc. Acad. Nat. Sci. Philadelphia, 114, 251–299. - get paper here
  • Malnate, Edmond V. 1960. Systematic division and evolution of the colubrid snake genus Natrix, with comments on the subfamily Natricinae. Proc. Acad. Nat. Sci. Philadelphia, 112 (3): 41-71. - get paper here
  • Pope, CLIFFORD H. 1935. The Reptiles of China. Turtes, Crocodilians, Snakes, Lizards. Amer. Mus. Nat. Hist., New York, Nat. Hist. Central Asia, 10: lii, 1-604 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Wu Lu, Li Te-chun, & Liu Chi-ch'en 1985. Kuei-chou p'a hsing lei chih [Reptiles of Kweichow Province]. [People's Press], Kuei-yang 436 pp.
  • Zhao E. M. 1998. Fauna Sinica, Reptilia, Vol. 3, Squamata: Serpentes. Science Press (In Chinese)
  • Zhao, E. (ed.) 2003. Coloured atlas of Sichuan reptiles [in Chinese]. Beijing, China Forestry Publishing House
  • Zhao,E. & Adler,K. 1993. Herpetology of China. SSAR, Oxford/Ohio, 1-522
 
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