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Dendrelaphis marenae VOGEL & VAN ROOIJEN, 2008

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Higher TaxaColubridae, Ahaetuliinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Maren’s Bronzeback 
SynonymDendrelaphis marenae VOGEL & VAN ROOIJEN 2008
Dendrelaphis marenae — GAULKE 2011: 276
Dendrelaphis marenae — DE LANG 2013
Dendrelaphis marenae — WALLACH et al. 2014: 218 
DistributionPhilippines [Balabac, Bantayan, Basilan, Bohol, Busuanga, Calauit (GAULKE 1999), Camiguin, Candaraman, Catanduanes (ROSS &GONZALES 1992), Cebu, Culion, Kalotkot, Leyte, Luzon, Masbate, Mindanao, Mindoro, Negros, Palawan, Polillo, Samar, Siquijor, Siargao (ROSS & LAZELL 1990), Sulu-Archipel (Bongao), Panay], Indonesia (Sulawesi).

Type locality: Province Albay, island of Luzon, Philippines.  
Reproductionoviparous. 
TypesHolotype: MNHN-RA 1994.0059, Coll. MONTAMO-RAY, date 1880 
DiagnosisDiagnosis – Aspecies of Dendrelaphis, characterized by the combination of: (1) vertebral scales enlarged, equal to or larger than the dorsals of the first row; (2) 159-191 ventrals; (3) 136-167 divided subcaudals; (4) 15 dorsal scale rows at midbody; (5) anal shield divided; (6) 1 loreal scale; (7) 2 or 3 supralabials touching the eye; (8) 9 (rarely 10) supralabials; (9) a long first sublabial that touches 4 infralabials on average (range: 2-5); (10) 4 to 8 temporals; (11) 2 (rarely 1 or 3) postoculars; (12) maximum total length 131.0 cm; (13) very long tail, TAIL/TL0.33-0.39; (14) a broad postocular stripe that starts on the postnasal or eye, covers the majority of the temporal region and extends onto the neck; (15) the presence of a light ventrolateral stripe and (16) black lines bordering the ventrolateral stripe absent or rudimentary. 
CommentDistribution: not on Panay (M. Gaulke, pers. comm., 26 Dec 2012).

Behavior: diurnal

Habitat: fully arboreal (Harrington et al. 2018) 
EtymologyThis species is named in honour of Dr. Maren GAULKE, for her contributions to the knowledge of the herpetology of the Philippines. She collected some of the paratypes and several other specimens used in this study. 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Blackburn, D.G. 1993. STANDARDIZED CRITERIA FOR THE RECOGNITION OF REPRODUCTIVE MODES IN SQUAMATE REPTILES. Herpetologica 49 (1): 118-132 - get paper here
  • Brown, R.M.; C.H. Oliveros; C.D. Siler; J.B. Fernandez; L.J. Welton; P.A.C. Buenavente; M.L.L. Diesmos; A.C. Diesmos 2012. Amphibians and Reptiles of Luzon Island (Philippines), VII: Herpetofauna of Ilocos Norte Province, Northern Cordillera Mountain Range. Check List 8(3):469-490 - get paper here
  • BROWN, RAFE M.; JOWARD B. BAUTISTA, MICHAEL A. CLORES & MICHAEL A. CUESTA. 2021. DENDRELAPHIS MARENAE (Gaulke’s Bronze-back Tree Snake). DIET. Herpetological Review 52 (1): 153–154. - get paper here
  • Brown; Rafe; Cameron Siler, Carl Oliveros, Luke Welton, Ashley Rock, John Swab, Merlijn Van Weerd, Jonah van Beijnen, Dominic Rodriguez, Edmund Jose, Arvin Diesmos 2013. The amphibians and reptiles of Luzon Island, Philippines, VIII: the herpetofauna of Cagayan and Isabela Provinces, northern Sierra Madre Mountain Range. ZooKeys 266 (2013) Special Issue: 1-120<br />doi: 10.3897/zookeys.266.3982 - get paper here
  • DEVAN-SONG, Anne and Rafe M. BROWN 2012. Amphibians and Reptiles of Luzon Island, Philippines, VI: The Herpetofauna of the Subic Bay Area. Asian Herpetological Research 3 (1): 1–20 - get paper here
  • Gaulke, M. 2011. The herpetofauna of Panay Island, Philippines. Edition Chimaira, 390 pp.
  • GLÄSSER-TROBISCH, A. & TROBISCH, D. 2020. Begegnungen mit Hydrosaurus & Co. in der Wallacea. Teil 1: Drachen im Reich der Schmetterlinge – Sulawesi Sauria 42 (3): 1-25 - get paper here
  • Gojo-Cruz, Paul Henric P. and Leticia E. Afuang 2018. The Zoogeographic Significance of Caraballo Mountain Range, Luzon Island, Philippines With Focus on the Biogeography of Luzon’s Herpetofauna. Philippine Journal of Science 147 (3): 393-409 - get paper here
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Koch, A. 2011. The Amphibians and Reptiles of Sulawesi: Underestimated Diversity in a Dynamic Environment. In: F.E. Zachos and J.C. Habel (eds.), Biodiversity Hotspots. Springer, Berlin, p. 383-404 - get paper here
  • Koch, A. 2012. Discovery, Diversity, and Distribution of the Amphibians and Reptiles of Sulawesi and its offshore islands. Edition Chimaira, 374 pp. [ISBN 978-3-89973-432-4] - get paper here
  • Lang, Ruud de 2013. The snakes of the Moluccas (Maluku), Indonesia. Edition Chimaira, 417 pp. - get paper here
  • Meneses CG, Siler CD, Alviola PA, Gonzalez JCT, Balatibat JB, Natividad CA, Brown RM 2022. Amphibian and reptile diversity along a ridge-to-reef elevational gradient on a small isolated oceanic island of the central Philippines. Check List 18(5): 941-948 - get paper here
  • Phangurha, J; Muhtianda, I A; Karya, A; Analuddin, K; Gillespie, G & Jones, S C; 2019. Dendrelaphis marenae (Maren’s Bronzeback) Predation behavior. Herpetological Review 50 (3): 590-591 - get paper here
  • Sanguila MB, Cobb KA, Siler CD, Diesmos AC, Alcala AC, Brown RM 2016. The amphibians and reptiles of Mindanao Island, southern Philippines, II: the herpetofauna of northeast Mindanao and adjacent islands. ZooKeys 624: 1–132, doi: 10.3897/zookeys.624.9814 - get paper here
  • Supsup CE, Asis AA, Carestia Jr UV, Diesmos AC, Mallari NAD, Brown RM 2020. Variation in species richness, composition and herpetological community structure across a tropical habitat gradient of Palawan Island, Philippines. Herpetozoa 33: 95-111 - get paper here
  • Supsup, Christian E.; Faith M. Guinto, Bernard R. Redoblado, Rai S. Gomez 2017. Amphibians and reptiles from the Mt. Hamiguitan Range of eastern Mindanao Island, Philippines: new distribution records. Check List 13 (3): 2121 - get paper here
  • SUPSUP, Christian E.; Nevong M. PUNA, Augusto A. ASIS, Bernard R. REDOBLADO, Maria Fatima G. PANAGUINIT, Faith M. GUINTO, Edmund B. RICO, Arvin C. DIESMOS, Rafe M. BROWN and Neil Aldrin D. MALLARI 2016. Amphibians and Reptiles of Cebu, Philippines: The Poorly Understood Herpetofauna of an Island with Very Little Remaining Natural Habitat. Asian Herpetological Research 2016, 7(3): 151–179 DOI: 10.16373/j.cnki.ahr.150049 - get paper here
  • Venturina, R. E. L., Diesmos, M. L. L., Maglangit, E. P. T., del Prado, Y. L. C., Ordas, J. A. D., Fernandez, J. B., ... & Diesmos, A. C. 2023. Herpetofauna of Mount Capistrano, a Fragmented Limestone Karst Forest in Central Mindanao, Philippines. Philippine Journal of Science, 152(6A), 2031-2048
  • Vogel, Gernot and Johan van Rooijen. 2008. Contributions to a review of the Dendrelaphis pictus (Gmelin, 1789) complex - 2. the eastern forms. Herpetozoa 21 (1/2): 3-29 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Weinell, Jeffrey L.; Errol Hooper, Alan E. Leviton, Rafe M. Brown 2019. Illustrated Key to the Snakes of the Philippines. Proc. Cal. Acad. Sci. (4) 66 (1): 1-49 - get paper here
 
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