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Bogertophis subocularis (BROWN, 1901)

IUCN Red List - Bogertophis subocularis - Least Concern, LC
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Higher TaxaColubridae, Colubrinae, Lampropeltini, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
SubspeciesBogertophis subocularis amplinotus WEBB 1990
Bogertophis subocularis subocularis (BROWN 1901) 
Common NamesE: subocularis: Trans-Pecos Rat Snake
G: Transpecos-Kletternatter, Transpecos-Rattennatter
S: Ratonera de Trans-Pecos 
SynonymColuber subocularis BROWN 1901: 492
Elaphe subocularis — STEJNEGER & BARBOUR 1917: 84
Elaphe sclerotica SMITH 1941 (nom. subst.)
Elaphe subocularis — AXTELL 1959
Elaphe subocularis — REYNOLDS & SCOTT 1977
Elaphe subocularis — WORTHINGTON 1981
Elaphe subocularis — NIEDERHAUSER 1984
Elaphe subocularis — STEBBINS 1985: 186
Elaphe subocularis — TANNER 1985: 630
Bogertophis subocularis — DOWLING & PRICE 1988
Bogertophis subocularis — CONANT & COLLINS 1991: 200
Bogertophis subocularis — LINER 1994
Bogertophis subocularis — CROTHER 2000: 57
Bogertophis subocularis — ERNST & ERNST 2003: 49
Bogertophis subocularis — CROTHER et al. 2012
Bogertophis subocularis — WALLACH et al. 2014: 100

Bogertophis subocularis amplinotus WEBB 1990
Elaphe subocularis amplinotus WEBB 1990

Bogertophis subocularis subocularis (BROWN 1901)
Coluber subocularis BROWN 1901
Bogertophis subocularis subocularis — CROTHER 2000: 57 
DistributionUSA (S New Mexico, W Texas),
Mexico (NE Chihuahua, Coahuila, NE Durango, Nuevo León)

amplinotus: Mexico (Durango, Coahuila); Type locality: Mexico: 90.5 mi N of City of Durango, Durango; R. Arndt and W. Sherbrooke; June 26, 1963.

subocularis: USA (W Texas, New Mexico), Mexico (Coahuila)

Type locality: USA: Texas, Jeff Davis County, Davis Mountains, 50 miles s-w from Pecos (Brown, 1901)  
Reproductionoviparous 
TypesType. ANSP 13733
Holotype: INHS (= UIMNH) 62817 [amplinotus] 
Diagnosis 
CommentSubspecies: Webb 1990 also described intergrades between subocularis and amplinotus from Coahuila and adjacent Nuevo Leon. B. s. amplinotus has been named for its larger body blotches as opposed to subocularis (Latin amplio = enlarge, expand; nota = mark).

Type species: Coluber subocularis BROWN 1901: 492 is the type species of the genus Bogertophis DOWLING & PRICE 1988.

Phylogenetics: see Dahn et al. 2018 for a phylogeny of Lampropeltini.

Similar species: B. rosaliae, 
References
  • Axtell, Ralph W. 1959. AMPHIBIANS AND REPTILES OF THE BLACK GAP WILDLIFE MANAGEMENT AREA, BREWSTER COUNTY, TEXAS. Southwestern Naturalist 4 (2): 88-109
  • Baker, R.J.; Bull, J.J.; Mengden, G.A. 1971. Chromosomes of Elaphe subocularis (Reptilia: Serpentes), with the description of an in vivo technique for preparation of snake chromosomes. Cellular and Molecular Life Sciences 27(10):1228-1229. - get paper here
  • Brown, A.E. 1901. A review of the Genera and Species of American Snakes, North of Mexico. Proc. Acad. Nat. Sci. Philad. 53: 10-110 - get paper here
  • Brown, A.E. 1902. A new species of Coluber from western Texas. Proceedings of the Academy of Natural Sciences of Philadelphia 53(3): 492–495 [1901] - get paper here
  • Campbell, Johnathan A. 1972. Reproduction in captive trans-pecos ratsnakes, Elaphe subocularis. Herpetological Review 4(4):129- 130. - get paper here
  • Conant,R. & Collins,J.T. 1991. A Field Guide to Reptiles and Amphibians of Eastern/Central North America, 3rd ed. Houghton Mifflin (Boston/New York), xx + 450 p.
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  • Dahn, H. A., Strickland, J. L., Osorio, A., Colston, T. J., & Parkinson, C. L. 2018. Hidden diversity within the depauperate genera of the snake tribe Lampropeltini (Serpentes, Colubridae). Molecular Phylogenetics and Evolution - get paper here
  • Davis DR, LaDuc TJ 2018. Amphibians and reptiles of C. E. Miller Ranch and the Sierra Vieja, Chihuahuan Desert, Texas, USA. ZooKeys 735: 97-130 - get paper here
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  • Dowling, H.G. and R. M. Price 1988. A proposed new genus for Elaphe subocularis and Elaphe rosaliae. The Snake 20: 52-63
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  • Lemos-Espinal JA, Smith GR, Cruz A 2016. Amphibians and Reptiles of the state of Nuevo León, Mexico. ZooKeys 594: 123-141, doi: 10.3897/zookeys.594.8289 - get paper here
  • Lemos-Espinal JA, Smith GR, Gadsden-Esparza H, Valdez-Lares R, Woolrich-Piña GA 2018. Amphibians and reptiles of the state of Durango, Mexico, with comparisons with adjoining states. ZooKeys 748: 65-87 - get paper here
  • Mattison, Chris 1995. The Encyclopedia of Snakes. New York: Facts on File, 256 pp.
  • Moll, E.O., and M. Rubio. 2006. Bogertophis subocularis (Brown, 1901), Trans-Pecos Ratsnake. Patronyms of the Pioneer West. Sonoran Herpetologist 19 (10):110-114. - get paper here
  • Nevárez-de-los-Reyes, Manuel, David Lazcano, Elí García-Padilla, Vicente Mata-Silva, Jerry D. Johnson and Larry David Wilson. 2016. The Herpetofauna of Nuevo León, Mexico: Composition, Distribution, and Conservation. Mesoamerican Herpetology 3 (3): 558–638 - get paper here
  • Niederhauser, R. 1984. Zuchterfolg bei der selten gehaltenen Transpecos-Natter, Elaphe subocularis. Herpetofauna 6 (31): 31-34 - get paper here
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  • Rhoads, D. D. 2016. Bogertophis subocularis (Trans-Pecos Ratsnake) diet and foraging behavior. Herpetological Review 47(3): 473. - get paper here
  • Rhoads, D. D. & Salmon, G. T. 2012. A much-belated obituary of an important American zoo collector, with discussion of the type locality for Bogertophis subocularis and Lampropeltis alterna. Herpetological Review 43: 270-273 - get paper here
  • Rhoads, Dustin D. 2016. Bogertophis subocularis (Trans-Pecos Ratsnake) mosaicism. Herpetological Review 47 (2): 241-242 - get paper here
  • Rhoads, Dustin D. 2017. Bogertophis subocularis (Trans-Pecos Ratsnake) Microhabitat Use. Herpetological Review 48 (1): 204 - get paper here
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  • Rocha, Arturo, Vicente Mata-Silva, Jerry D. Johnson and William D. Lukefahr. 2013. Bogertophis subocularis (Trans-Pecos ratsnake) overwintering behavior. Herpetological Review 44 (3): 517-518 - get paper here
  • Schulz, K. D. 1996. Eine Monographie der Schlangengattung Elaphe Fitzinger. Bushmaster, Berg (CH): 1-460
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  • Webb, R. G. 1990. Description of a new subspecies of Bogertophis subocularis (Brown) from northern Mexico (Serpentes: Colubridae). Texas Journal of Science 42 (3): 227-243 - get paper here
  • Webb, Robert G.;Ferguson, George M. 1986. Morphological variation in the trans-Pecos rat snake (Elaphe subocularis). Southwestern Naturalist 31 (1): 118-121 - get paper here
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