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Atractus pantostictus FERNANDES & PUORTO, 1993

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesPortuguese: Cobra-da-Terra, Fura-Terra 
SynonymAtractus pantostictus FERNANDES & PUORTO 1993
Atractus guentheri – PETERS & OREJAS-MIRANDA 1970:29. (part)
Atractus pantostictus — PASSOS et al. 2005
Atractus pantostictus — PASSOS et al. 2007
Atractus pantostictus — WALLACH et al. 2014: 78
Atractus pantostictus — NOGUEIRA et al. 2019 
DistributionBrazil (Tocantins, São Paulo)

Type locality: municipality of Franco da Rocha (23º20’S, 46º43’W, ca. 800 m), state of São Paulo, Brazil  
Reproductionoviparous 
TypesHolotype: IBSP (= IB) 54844 
DiagnosisDiagnosis: Atractus pantostictus is distinguished from all congeners by the following combination of characters: (1) 17 smooth dorsal scale rows at midbody; (2) two postoculars; (3) moderate loreal; (4) temporals 1+2; (5) seven supralabials, third and fourth contacting chinshields; (6) usually seven infralabials, first four contacting chinshields; (7) six to eight maxillary teeth; (8) three gular scale rows; (9) usually three preventrals; (10) 154–170 ventrals in females, 145–166 in males; (11) 19–28 subcaudals in females, 22–33 in males; (12) dorsal ground colour creamish yellow to brown uniformly scattered with small black dots; (13) venter immaculate creamish white; (14) moderate body size, females reaching 390 mm SVL, males 322 mm SVL; (15) tail short to moderate in females (6.4–13.2% SVL) and short to long (8.0–17.6% SVL) in males; (16) hemipenis slightly bilobed, slightly semicapitate, and semicalyculate [from PASSOS et al. 2010]. 
CommentCan be confused with A. guentheri. 
References
  • BARBO, F. E.; MARQUES, O. A. V. & SAWAYA, R. J. 2011. Diversity, Natural History and Distribution of Snakes in the Municipality of São Paulo. South American J. Herp. 6: 135-160
  • Cacciali, P.; Villalba, R. & Yanosky, A.A. 2007. A NEW SPECIES OF ATRACTUS (SERPENTES:COLUBRIDAE: DIPSADINAE) FROM THE ATLANTIC FOREST OF ALTO PARANÁ, PARAGUAY. South American J. Herp. 2 (2): 83-88 - get paper here
  • Fernandes R. & Puorto G. 1993. A new species of Atractus from Brazil and the status of A. guentheri (Serpentes: Colubridae). Mem. Inst. Butantan 55 (SUPPL. 1): 7-14. - get paper here
  • Fiorillo, B. F., Maciel, J. H., & Martins, M. 2021. Composition and natural history of a snake community from the southern Cerrado, southeastern Brazil. ZooKeys 1056: 95-147 - get paper here
  • Franca, F.G.R.; Daniel O. Mesquita, Cristiano C. Nogueira, and Alexandre F. B. Araújo 2008. Phylogeny and Ecology Determine Morphological Structure in a Snake Assemblage in the Central Brazilian Cerrado. Copeia 2008 (1): 23-38 - get paper here
  • Gonzalez R. C. et al. 2020. Lista dos Nomes Populares dos Répteis no Brasil – Primeira Versão. Herpetologia Brasileira 9 (2): 121 – 214 - get paper here
  • Guerra, V., Ramalho, W. P., Machado, I. F., & Brandão, R. A. 2022. Herpetofauna of the Serra do Tombador Nature Reserve, State of Goiás, Central Brazil. Arquivos de Zoologia, 53(3), 33-51 - get paper here
  • MARQUES, O. A. V.; PEREIRA, D. N.; BARBO, F.E.; GERMANO, V.J. & SAWAYA, R. J. 2009. Os Répteis do Município de São Paulo: diversidade e ecologia da fauna pretérita e atual. Biota Neotropica 9: 1-12 - get paper here
  • Nogueira, Cristiano C.; Antonio J.S. Argôlo, Vanesa Arzamendia, Josué A. Azevedo, Fausto E. Barbo, Renato S. Bérnils, Bruna E. Bolochio, Marcio Borges-Martins, Marcela Brasil-Godinho, Henrique Braz, Marcus A. Buononato, Diego F. Cisneros-Heredia, 2019. Atlas of Brazilian snakes: verified point-locality maps to mitigate the Wallacean shortfall in a megadiverse snake fauna. South American J. Herp. 14 (Special Issue 1):1-274 - get paper here
  • Passos, P., R. Fernandes and Borges-Nojosa, D.M. 2007. A New Species of Atractus (Serpentes: Dipsadinae) from a Relictual Forest in Northeastern Brazil. Copeia 2007 (4): 788–797 - get paper here
  • Passos, P., R. Fernandes and N. Zanella 2005. A new species of Atractus (Serpentes: Colubridae) from Southern Brazil. Herpetologica 61 (2): 209-218 - get paper here
  • Passos, P.; Fernandes, R.; Bernils, R.S. & Moura-Leite, J.C. de 2010. Taxonomic revision of the Brazilian Atlantic Forest Atractus (Reptilia: Serpentes: Dipsadidae). Zootaxa 2364: 1–63 - get paper here
  • RECODER, R.S.,TEIXEIRA JUNIOR, M., CAMACHO, A., NUNES, P.M.S., MOTT, T., VALDUJO, P.H., GHELLERE, J.M., NOGUEIRA, C. & RODRIGUES, M.T. 2011. Reptiles of Serra Geral do Tocantins Ecological Station, Central Brazil. Biota Neotrop. 11(1): - get paper here
  • RESENDE, FLÁVIA CAPPUCCIO & LUCIANA BARRETO NASCIMENTO. 2022. Sexual Segment of the Kidney and Testicular Activity of the Neotropical Snake Atractus pantostictus (Squamata: Dipsadidae) from Minas Gerais State, Southeastern Brazil. South American Journal of Herpetology 24(1): 11–18. - get paper here
  • Sawaya, Ricardo Jannini; Otavio Augusto Vuolo Marques <br />& Marcio Martins 2008. Composition and natural history of a Cerrado snake assemblage at Itirapina, São Paulo state, southeastern Brazil Biota Neotropica 8 (2): 127-149 - get paper here
  • SOUSA, B.M., NASCIMENTO, A.E.R., GOMIDES, S.C., VARELA RIOS, C.H., HUDSON, A.H. & NOVELLI, I.A. 2010. Reptiles in fragments of Cerrado and Atlantic Forest at the Campo das Vertentes, Minas Gerais State, Southeastern Brazil. Biota Neotropica 10 (2): 129-138 - get paper here
  • Torello-Viera, Natália F. and Otavio A.V. Marques 2017. Daily Activity of Neotropical Dipsadid Snakes. South American J. Herp. 12 (2): 128-135. - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Zaher, Hussam; Fausto Erritto BarboI; Paola Sanchez Martínez; Cristiano Nogueira; Miguel Trefaut Rodrigues; Ricardo Jannini Sawaya 2011. Répteis do Estado de São Paulo: conhecimento atual e perspectivas. Biota Neotropica, 11 (1): 1–15. - get paper here
 
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